Catalogue of the Orphninae
Tribe ORPHNINI Erichson, 1847
Type genus: Orphnus MacLeay, 1819
Chaetonyx Schaum, 1862
Type species: Chaetonyx robustus Schaum, by monotypy.
- Chaetonyx binaghii Mariani, 1946
-
Chaetonyx robustus Schaum, 1862
ssp. Chaetonyx robustus italicus Mariani, 1946
ssp. Chaetonyx robustus liguricus Mariani, 1946
ssp. Chaetonyx robustus robustus Schaum, 1862 - Chaetonyx schatzmayri Mariani, 1946
Hybalus Dejean, 1833
syn. Geobius Brullé, 1832
Type species: Geobius cornifrons Brullé, by monotypy.
- Hybalus algiricus Petrovitz, 1968
- Hybalus alonsoi López-Colón, 2014
-
Hybalus ameliae López-Colón, 1986
syn. Hybalus ameliae baguenae López-Colón, 1986 - Hybalus angustatus Lucas, 1855
- Hybalus arenicola Baraud, 1991
- Hybalus atlanticus López-Colón, 1992
- Hybalus barbarus (Laporte de Castelnau, 1840)
- Hybalus baudoni Petrovitz, 1964
- Hybalus benoitii Tournier, 1864
- Hybalus bigibber Reitter, 1893
- Hybalus bletoni Baraud 1991
- Hybalus constantini Baraud, 1979
-
Hybalus cornifrons (Brullé, 1832)
syn. Hybalus graecus Sturm, 1843
syn. Hybalus dorcas Germar, 1838 - Hybalus digitatus Petrovitz, 1963
-
Hybalus dorcas (Fabricius, 1798)
syn. Geobius tingitanus Fairmaire, 1852
syn. Hybalus reclinans Fairmaire, 1879
syn. Hybalus quedenfeldti Petrovitz, 1964 -
Hybalus doursii Lucas, 1853
syn. Hybalus gazella Raffray in Fairmaire et Raffray, 1873
syn. Hybalus raffrayi Arrow, 1911 -
Hybalus glabratus (Fabricius, 1792)
syn. Hybalus cornifrons Guérin-Méneville, 1844 -
Hybalus granicornis Fairmaire, 1877
syn. Hybalus biretusus Marseul, 1878 - Hybalus kocheri Petrovitz, 1964
- Hybalus maroccanus Petrovitz, 1964
- Hybalus normandi Baraud, 1980
- Hybalus numidicus Petrovitz, 1964
- Hybalus parvicornis Lucas, 1855
- Hybalus petrovitzi Baraud, 1991
- Hybalus punicus Baraud, 1991
- Hybalus pygmaeus (Quensel, 1806)
- Hybalus ramicornis Reitter, 1893
- Hybalus reflexus Petrovitz, 1964
-
Hybalus rotroui Petrovitz, 1964
ssp. Hybalus rotroui rotroui Petrovitz, 1964
ssp. Hybalus rotroui peyerimhoffi López-Colón, 1991 - Hybalus saezi López-Colón, 1992
-
Hybalus servulus Normand, 1949
syn. Hybalus demoflysi Baraud, 1980 - Hybalus subcornutus Fairmaire, 1870
- Hybalus sulcatus Baraud 1991
- Hybalus tricornis (Lucas, 1849)
- Hybalus tuberculicornis Reitter, 1893
- Hybalus varians Petrovitz, 1964
Род Orphnus Macleay, 1819
Type species: Scarabaeus bicolor Fabricius, 1801, by monotypy.
Subgenus Cerhomalus Quedenfeldt, 1884
Type species: Orphnus mechowi (Quedenfeldt, 1884), by monotypy.
- Orphnus absconditus Petrovitz, 1971
- Orphnus mechowi (Quedenfeldt, 1884)
Subgenus Horpnus Paulian, 1948
Type species: Orphnus ellenbergeri Paulian, 1948, by monotypy.
- Orphnus asperatus Petrovitz, 1971
- Orphnus ellenbergeri Paulian, 1948
- Orphnus zumpti Petrovitz, 1963
Subgenus Orphnus s. str. Macleay, 1819
Type species: bicolor (Fabricius, 1801), by monotypy.
= Subgenus Parorphnus Paulian, 1948
- Orphnus acuticornis Petrovitz, 1971
- Orphnus amplitarsis Petrovitz, 1971
- Orphnus angolensis Quedenfeldt, 1884
- Orphnus babaulti Paulian, 1948
- Orphnus benderitteri Pic, 1930
- Orphnus bicolor (Fabricius, 1801)
- Orphnus bifidus Schmidt, 1912
-
Orphnus bilobus Klug, 1855
syn. Orphnus mashunus Péringuey, 1896 - Orphnus brunneus Benderitter, 1912
- Orphnus camerunensis Petrovitz, 1971
- Orphnus capensis Petrovitz, 1971
- Orphnus chappuisi Paulian, 1951
- Orphnus compactilis Quedenfeldt, 1884
- Orphnus compactus Petrovitz, 1971
- Orphnus compressicornis Benderitter, 1913
- Orphnus congolanus Petrovitz, 1971
- Orphnus convexus Benderitter, 1913
- Orphnus copridoides Paulian, 1948
- Orphnus costatus Petrovitz, 1971
- Orphnus crassus Pic, 1928
- Orphnus cribratellus Fairmaire, 1878
-
Orphnus declivis Schmidt, 1912
ssp. Orphnus declivis declivis Schmidt, 1912
ssp. Orphnus declivis baloghi Petrovitz, 1971 - Orphnus drumonti Frolov, 2009
- Orphnus fossatus Paulian, 1951
- Orphnus foveolatus Benderitter, 1920
- Orphnus galla Gestro, 1895
- Orphnus gilleti Benderitter, 1923
- Orphnus grossepunctatus Petrovitz, 1971
- Orphnus guineensis Petrovitz, 1971
- Orphnus harrisoni Frolov, 2009
- Orphnus herero Petrovitz, 1963
- Orphnus heteronychoides Paulian, 1948
- Orphnus imitator Benderitter, 1920
- Orphnus impressus Westwood, 1846
-
Orphnus incultus (Péringuey, 1892)
syn. Hybosorus incultus Péringuey, 1892 - Orphnus jeanneli Benderitter, 1914
- Orphnus kafuenus Péringuey, 1908
- Orphnus leleupi Petrovitz, 1971
- Orphnus letestui Paulian, 1948
- Orphnus longicornis Frolov & Akhmetova, 2015
- Orphnus luluanus Paulian, 1948
- Orphnus luminosus Benderitter, 1920
-
Orphnus macleayi Laporte de Castelnau, 1832
syn. Orphnus arrowi Benderitter, 1912
syn. Orphnus meleagris Westwood, 1846
ssp. Orphnus macleayi macleayi Laporte de Castelnau, 1832
ssp. Orphnus macleayi zambezianus Péringuey, 1896
syn. Orphnus meleagris confluens Benderitter, 1920
syn. Orphnus meleagris latus Benderitter, 1920
ssp. Orphnus macleayi emeritus Péringuey, 1901 - Orphnus mandibularis (Lansberge, 1886)
- Orphnus mashunensis Péringuey, 1908
- Orphnus mombasaensis Benderitter, 1914
- Orphnus mpese Paulian, 1948
- Orphnus mysoriensis Westwood, 1846
- Orphnus niger Pic, 1928
- Orphnus nyassicus Kolbe, 1895
-
Orphnus orbus Benderitter, 1920
ssp. Orphnus orbus orbus Benderitter, 1920
ssp. Orphnus orbus orientalis Paulian 1948 - Orphnus oryctoides Quedenfeldt, 1888
- Orphnus ovampoanus Péringuey, 1896
- Orphnus overlaeti Petrovitz, 1971
- Orphnus parentalis Péringuey, 1908
-
Orphnus parvus (Wiedemann, 1823)
syn. Orphnus nanus Westwood, 1846 - Orphnus pauliani Gomes Alves, 1957
- Orphnus pici Paulian, 1948
- Orphnus picinus Westwood, 1846
- Orphnus plebejus Péringuey, 1901
- Orphnus posthi Paulian, 1948
- Orphnus pugnax Péringuey, 1896
- Orphnus rufithorax Benderitter, 1914
- Orphnus rufulus Boheman, 1857
- Orphnus sansibaricus Kolbe, 1895
- Orphnus schoutedeni Benderitter, 1920
- Orphnus senegalensis Laporte de Castelnau, 1832
-
Orphnus similis Petrovitz, 1971
ssp. Orphnus similis similis Petrovitz, 1971
ssp. Orphnus similis umbilicatus Petrovitz, 1971 - Orphnus sinuaticeps Petrovitz, 1971
- Orphnus sinuatus Benderitter, 1923
- Orphnus striatoides Paulian, 1948
-
Orphnus striatopunctatus Felsche, 1904
syn. Orphnus clypeatus Benderitter, 1920
syn. Orphnus felschei Schmidt, 1912 - Orphnus striatus Benderitter, 1913
- Orphnus subcornutus Paulian, 1948
- Orphnus subfurcatus Kolbe, 1895
- Orphnus sudanicus Paulian, 1948
- Orphnus tchadensis Benderitter, 1913
- Orphnus thoracicus Linell, 1896
- Orphnus tinantae Paulian, 1948
- Orphnus transvaalensis Frolov, 2009
- Orphnus tristis Pic, 1928
- Orphnus usambaricus Petrovitz, 1971
- Orphnus viduae Petrovitz, 1971
Subgenus Pachyorphnus Paulian, 1948
Type species: Orphnus subfoveatus Fairmaire, 1898, by original designation.
- Orphnus clavipes Petrovitz, 1971
- Orphnus dewittei Petrovitz, 1971
- Orphnus subfoveatus Fairmaire, 1898
Subgenus Phornus Paulian, 1948
Type species: Orphnus strangulatus Paulian, 1948, by original designation.
- Orphnus ferrierei Frolov et Akhmetova, 2016
- Orphnus giganteus Paulian, 1948
- Orphnus parastrangulatus Frolov et Akhmetova, 2016
- Orphnus strangulatus Paulian, 1948
- Orphnus renaudi Frolov et Akhmetova, 2016
- Orphnus valeriae Frolov et Akhmetova, 2016
Subgenus Ronphus Paulian, 1948
Type species: Orphnus livingstonei Paulian, 1948, by original designation.
- Orphnus demeyeri Frolov, 2013
- Orphnus livingstonei Paulian, 1948
- Orphnus massarti Paulian, 1948
- Orphnus peringueyi Paulian, 1948
- Orphnus planicollis Petrovitz, 1971
- Orphnus quadrigibbosus Petrovitz, 1971
- Orphnus simonii Petrovitz, 1971
- Orphnus testaceus Paulian, 1948
Craniorphnus Kolbe 1895
Type species: Craniorphnus grandiceps Kolbe, 1895, by monotypy.
- Craniorphnus grandiceps Kolbe, 1895
Goniorphnus Arrow, 1911
Type species: Goniorphnus felschei Arrow, 1911, by monotypy.
- Goniorphnus felschei Arrow, 1911
Hybaloides Quedenfeldt, 1884
Type species: Hybaloides foveolatus Quedenfeldt, 1884, by monotypy.
- Hybaloides foveolatus Quedenfeldt, 1884
Pseudorphnus Benderitter, 1913
Type species: Orphnus coquerelii Fairmaire, 1868, by monotypy.
- Pseudorphnus carinatus Frolov, 2011
- Pseudorphnus coquerelii (Fairmaire, 1868)
- Pseudorphnus hiboni Paulian, 1959
- Pseudorphnus olsoufieffi Paulian, 1977
Madecorphnus Paulian, 1992
Type species: Drepanognathus falciger Lansberge, 1886, original designation.
- Madecorphnus aquilonius Frolov, 2012
- Madecorphnus barclayi Frolov, 2012
- Madecorphnus brunneus Frolov, 2010
- Madecorphnus cuccodoroi Frolov, 2011
- Madecorphnus dentatus Frolov, 2010
- Madecorphnus falcatus Paulian, 1992
- Madecorphnus falciger (Lansberge, 1886)
- Madecorphnus falculoides (Paulian, 1977)
- Madecorphnus fisheri Frolov, 2014
- Madecorphnus grandis Frolov, Smith et Akhmetova, 2017
- Madecorphnus hanskii Frolov, 2012
- Madecorphnus montreuili Frolov, 2010
- Madecorphnus niger Frolov, 2010
- Madecorphnus pauliani Frolov, 2010
- Madecorphnus perinetensis Frolov, 2010
- Madecorphnus peyrierasi Frolov, 2010
- Madecorphnus punctatus Frolov, 2010
- Madecorphnus saintemariensis Frolov, 2014
- Madecorphnus simplex Frolov, 2010
- Madecorphnus tuberculatus Frolov, 2012
Triodontus Westwood, 1845
syn. Orphnidius (Benderitter, 1913)
Type species: Orphnus nitidulus Guérin-Méneville, 1844, by monotypy.
-
Triodontus alticola Paulian, 1977
syn. Triodontus maroantsetrae Paulian, 1977. Synonymy by Frolov et al. (2016) - Triodontus ankarafantsikae Frolov, Montreuil et Akhmetova, 2016
- Triodontus bicavatus (Fairmaire, 1905)
- Triodontus copridoides Paulian, 1977
- Triodontus fairmairei Frolov, Montreuil et Akhmetova, 2016
- Triodontus hanskii Frolov, 2010
- Triodontus hildebrandtii (Fairmaire, 1883)
-
Triodontus hova (Fairmaire, 1868)
syn. Orphnus nigrita Brancsik, 1893. Synonymy by Frolov et al. (2016) - Triodontus inexpectatus Frolov, Montreuil et Akhmetova, 2016
- Triodontus itremoi Paulian, 1977
- Triodontus lemoulti Frolov, Montreuil et Akhmetova, 2016
- Triodontus modestus (Benderitter, 1914)
-
Triodontus nitidulus (Guérin-Méneville, 1844)
syn. Orphnus obsoletus Brancsik, 1893. Synonymy by Frolov et al. (2016)
syn. Triodontus occidentalis Paulian, 1977. Synonymy by Frolov et al. (2016) -
Triodontus owas Westwood, 1852
syn. Triodontus vadoni Paulian, 1977. Synonymy by Frolov et al. (2016)
syn. Triodontus perrotorum Paulian, 1977. Synonymy by Frolov et al. (2016) - Triodontus viettei Frolov, Montreuil et Akhmetova, 2016
Renorphnus Frolov et Montreuil, 2009
Type species: Orphnus clementi Petrovitz, 1971, by monotypy.
- Renorphnus clementi (Petrovitz, 1971)
Tribe AEGIDIINI Paulian, 1984
Type genus: Aegidium Westwood, 1845
Aegidium Westwood, 1845
Type species: Aegidium colombianum Westwood.
- Aegidium atlanticum Frolov, Grossi, & Vaz-de-Mello, 2015
- Aegidium angustum Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium bolivianum Frolov & Akhmetova, 2020
- Aegidium bordoni Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium borrei Paulian, 1984
- Aegidium chalumeaui Rojkoff et Frolov, 2017
-
Aegidium colombianum Westwood, 1845
syn. Aegidium elongatum Paulian, 1984, synonymy by Frolov et al. (2017). -
Aegidium cribratum Bates, 1887
syn. Aegidium asperatum Preudhomme de Borre, 1886, synonymy by Frolov et al. (2017).
syn. Aegidium squamatum Bates, 1887: synonymy by Frolov et al. (2017). - Aegidium dierkensi Rojkoff et Frolov, 2017
- Aegidium dominicense Cartwright and Chalumeau, 1977
- Aegidium geayi Paulian, 1984
- Aegidium gilli Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium hostilioi Gasca-Álvarez and Deloya, 2020
- Aegidium howdeni Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium minor Paulian, 1984
- Aegidium parvulus Westwood 1845
- Aegidium onorei Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium peresfilhoi Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium peruanum Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium rafaeli Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium reichei Preudhomme de Borre, 1886
- Aegidium sinuatum Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium varians Frolov, Akhmetova & Vaz-de-Mello, 2017
- Aegidium vincentiae Arrow, 1903
Aegidiellus Paulian, 1984
Type species: Phileurus alatus Laporte de Castelnau, by monotypy.
-
Aegidiellus alatus (Laporte de Castelnau, 1840)
syn. Aegidium haedulus Westwood, 1845 - Aegidiellus dentipenis Frolov, Akhmetova et Vaz-de-Mello, 2017
- Aegidiellus zezaoi Frolov, Akhmetova et Vaz-de-Mello, 2017
Aegidinus Arrow, 1904
Type species: Aegidium guianensis Westwood, designated by Paulian (1984).
- Aegidinus brasiliensis Arrow, 1904
-
Aegidinus candezei (Preudhomme de Borre, 1886)
syn. Aegidinus sunidigea Colby, 2009 - Aegidinus colbyae Frolov, Akhmetova & Vaz-de-Mello, 2019
-
Aegidinus cornutus Colby, 2009
syn. Aegidinus unicus Colby, 2009 -
Aegidinus guianensis (Westwood, 1845)
syn. Aegidinus crypticus Colby, 2009
syn. Aegidinus oreibates Colby, 2009 - Aegidinus howdenorum Colby, 2009
- Aegidinus howeae Colby, 2009
- Aegidinus noriegai Frolov, Akhmetova & Vaz-de-Mello, 2019
- Aegidinus petrovi Colby, 2009
- Aegidinus simulatus Colby, 2009
- Aegidinus teamscaraborum Colby, 2009
- Aegidinus steinheili (Harold, 1880)
- Aegidinus tricornis Colby, 2009
- Aegidinus venezuelensis Colby, 2009
Onorius Frolov et Vaz-de-Mello, 2015
Type species: Onorius inexpectatus Frolov et Vaz-de-Mello, 2015, by monotypy.
Paraegidium Vulcano, Pereira, et Martínez, 1966
Type species: Paraegidium costalimai Vulcano, Pereira, et Martínez, by monotypy.
- Paraegidium barretoi Frolov, Akhmetova & Vaz-de-Mello, 2017
- Paraegidium costalimai Vulcano, Pereira, et Martínez, 1966
- Paraegidium howdeni Frolov, Akhmetova & Vaz-de-Mello, 2017
- Paraegidium monneorum Frolov, Akhmetova & Vaz-de-Mello, 2017
- Paraegidium oliveirai Frolov, Akhmetova & Vaz-de-Mello, 2017
- Paraegidium pokornyi Frolov, Akhmetova & Vaz-de-Mello, 2017
Orphninae incertae sedis
Stenosternus Karsch, 1881
Type species: Stenosternus costatus Karsch, 1881, by monotypy.
- Stenosternus costatus Karsch, 1881