Catalogue of the Orphninae

Tribe ORPHNINI Erichson, 1847

Type genus: Orphnus MacLeay, 1819

Chaetonyx Schaum, 1862

Type species: Chaetonyx robustus Schaum, by monotypy.

1. Chaetonyx binaghii Mariani, 1946
2. Chaetonyx robustus Schaum, 1862
    ssp. Chaetonyx robustus italicus Mariani, 1946
    ssp. Chaetonyx robustus liguricus Mariani, 1946
    ssp. Chaetonyx robustus robustus Schaum, 1862
3. Chaetonyx schatzmayri Mariani, 1946

Hybalus Dejean, 1833

syn. Geobius Brullé, 1832
Type species: Geobius cornifrons Brullé, by monotypy.

1. Hybalus algiricus Petrovitz, 1968
2. Hybalus alonsoi López-Colón, 2014
3. Hybalus ameliae López-Colón, 1986
    syn. Hybalus ameliae baguenae López-Colón, 1986
4. Hybalus angustatus Lucas, 1855
5. Hybalus arenicola Baraud, 1991
6. Hybalus atlanticus López-Colón, 1992
7. Hybalus barbarus (Laporte de Castelnau, 1840)
8. Hybalus baudoni Petrovitz, 1964
9. Hybalus benoitii Tournier, 1864
10. Hybalus bigibber Reitter, 1893
11. Hybalus bletoni Baraud 1991
12. Hybalus constantini Baraud, 1979
13. Hybalus cornifrons (Brullé, 1832)
     syn. Hybalus graecus Sturm, 1843
     syn. Hybalus dorcas Germar, 1838
14. Hybalus digitatus Petrovitz, 1963
15. Hybalus dorcas (Fabricius, 1798)
     syn. Geobius tingitanus Fairmaire, 1852
     syn. Hybalus reclinans Fairmaire, 1879
     syn. Hybalus quedenfeldti Petrovitz, 1964
16. Hybalus doursii Lucas, 1853
     syn. Hybalus gazella Raffray in Fairmaire et Raffray, 1873
     syn. Hybalus raffrayi Arrow, 1911
17. Hybalus glabratus (Fabricius, 1792)
     syn. Hybalus cornifrons Guérin-Méneville, 1844
18. Hybalus granicornis Fairmaire, 1877
     syn. Hybalus biretusus Marseul, 1878
19. Hybalus kocheri Petrovitz, 1964
20. Hybalus maroccanus Petrovitz, 1964
21. Hybalus normandi Baraud, 1980
22. Hybalus numidicus Petrovitz, 1964
23. Hybalus parvicornis Lucas, 1855
24. Hybalus petrovitzi Baraud, 1991
25. Hybalus punicus Baraud, 1991
26. Hybalus pygmaeus (Quensel, 1806)
27. Hybalus ramicornis Reitter, 1893
28. Hybalus reflexus Petrovitz, 1964
29. Hybalus rotroui Petrovitz, 1964
     ssp. Hybalus rotroui rotroui Petrovitz, 1964
     ssp. Hybalus rotroui peyerimhoffi López-Colón, 1991
30. Hybalus saezi López-Colón, 1992
31. Hybalus servulus Normand, 1949
     syn. Hybalus demoflysi Baraud, 1980
32. Hybalus subcornutus Fairmaire, 1870
33. Hybalus sulcatus Baraud 1991
34. Hybalus tricornis (Lucas, 1849)
35. Hybalus tuberculicornis Reitter, 1893
36. Hybalus varians Petrovitz, 1964

Род Orphnus Macleay, 1819

Type species: Scarabaeus bicolor Fabricius, 1801, by monotypy.

Subgenus Cerhomalus Quedenfeldt, 1884
Type species: Orphnus mechowi (Quedenfeldt, 1884), by monotypy.

1. Orphnus absconditus Petrovitz, 1971
2. Orphnus mechowi (Quedenfeldt, 1884)

Subgenus Horpnus Paulian, 1948
Type species: Orphnus ellenbergeri Paulian, 1948, by monotypy.

3. Orphnus asperatus Petrovitz, 1971
4. Orphnus ellenbergeri Paulian, 1948
5. Orphnus zumpti Petrovitz, 1963

Subgenus Orphnus s. str. Macleay, 1819
Type species: bicolor (Fabricius, 1801), by monotypy.
= Subgenus Parorphnus Paulian, 1948

6. Orphnus acuticornis Petrovitz, 1971
7. Orphnus amplitarsis Petrovitz, 1971
8. Orphnus angolensis Quedenfeldt, 1884
9. Orphnus babaulti Paulian, 1948
10. Orphnus benderitteri Pic, 1930
11. Orphnus bicolor (Fabricius, 1801)
12. Orphnus bifidus Schmidt, 1912
13. Orphnus bilobus Klug, 1855
     syn. Orphnus mashunus Péringuey, 1896
14. Orphnus brunneus Benderitter, 1912
15. Orphnus camerunensis Petrovitz, 1971
16. Orphnus capensis Petrovitz, 1971
17. Orphnus chappuisi Paulian, 1951
18. Orphnus compactilis Quedenfeldt, 1884
19. Orphnus compactus Petrovitz, 1971
20. Orphnus compressicornis Benderitter, 1913
21. Orphnus congolanus Petrovitz, 1971
22. Orphnus convexus Benderitter, 1913
23. Orphnus copridoides Paulian, 1948
24. Orphnus costatus Petrovitz, 1971
25. Orphnus crassus Pic, 1928
26. Orphnus cribratellus Fairmaire, 1878
27. Orphnus declivis Schmidt, 1912
     ssp. Orphnus declivis declivis Schmidt, 1912
     ssp. Orphnus declivis baloghi Petrovitz, 1971
28. Orphnus drumonti Frolov, 2009
29. Orphnus fossatus Paulian, 1951
30. Orphnus foveolatus Benderitter, 1920
31. Orphnus galla Gestro, 1895
32. Orphnus gilleti Benderitter, 1923
33. Orphnus grossepunctatus Petrovitz, 1971
34. Orphnus guineensis Petrovitz, 1971
35. Orphnus harrisoni Frolov, 2009
36. Orphnus herero Petrovitz, 1963
37. Orphnus heteronychoides Paulian, 1948
38. Orphnus imitator Benderitter, 1920
39. Orphnus impressus Westwood, 1846
40. Orphnus incultus (Péringuey, 1892)
     syn. Hybosorus incultus Péringuey, 1892
41. Orphnus jeanneli Benderitter, 1914
42. Orphnus kafuenus Péringuey, 1908
43. Orphnus leleupi Petrovitz, 1971
44. Orphnus letestui Paulian, 1948
45. Orphnus longicornis Frolov & Akhmetova, 2015
46. Orphnus luluanus Paulian, 1948
47. Orphnus luminosus Benderitter, 1920
48. Orphnus macleayi Laporte de Castelnau, 1832
     syn. Orphnus arrowi Benderitter, 1912
     syn. Orphnus meleagris Westwood, 1846
     ssp. Orphnus macleayi macleayi Laporte de Castelnau, 1832
     ssp. Orphnus macleayi zambezianus Péringuey, 1896
     syn. Orphnus meleagris confluens Benderitter, 1920
     syn. Orphnus meleagris latus Benderitter, 1920
     ssp. Orphnus macleayi emeritus Péringuey, 1901
49. Orphnus mandibularis (Lansberge, 1886)
50. Orphnus mashunensis Péringuey, 1908
51. Orphnus mombasaensis Benderitter, 1914
52. Orphnus mpese Paulian, 1948
53. Orphnus mysoriensis Westwood, 1846
54. Orphnus niger Pic, 1928
55. Orphnus nyassicus Kolbe, 1895
56. Orphnus orbus Benderitter, 1920
     ssp. Orphnus orbus orbus Benderitter, 1920
     ssp. Orphnus orbus orientalis Paulian 1948
57. Orphnus oryctoides Quedenfeldt, 1888
58. Orphnus ovampoanus Péringuey, 1896
59. Orphnus overlaeti Petrovitz, 1971
60. Orphnus parentalis Péringuey, 1908
61. Orphnus parvus (Wiedemann, 1823)
     syn. Orphnus nanus Westwood, 1846
62. Orphnus pauliani Gomes Alves, 1957
63. Orphnus pici Paulian, 1948
64. Orphnus picinus Westwood, 1846
65. Orphnus plebejus Péringuey, 1901
66. Orphnus posthi Paulian, 1948
67. Orphnus pugnax Péringuey, 1896
68. Orphnus rufithorax Benderitter, 1914
69. Orphnus rufulus Boheman, 1857
70. Orphnus sansibaricus Kolbe, 1895
71. Orphnus schoutedeni Benderitter, 1920
72. Orphnus senegalensis Laporte de Castelnau, 1832
73. Orphnus similis Petrovitz, 1971
     ssp. Orphnus similis similis Petrovitz, 1971
     ssp. Orphnus similis umbilicatus Petrovitz, 1971
74. Orphnus sinuaticeps Petrovitz, 1971
75. Orphnus sinuatus Benderitter, 1923
76. Orphnus striatoides Paulian, 1948
77. Orphnus striatopunctatus Felsche, 1904
     syn. Orphnus clypeatus Benderitter, 1920
     syn. Orphnus felschei Schmidt, 1912
78. Orphnus striatus Benderitter, 1913
79. Orphnus subcornutus Paulian, 1948
80. Orphnus subfurcatus Kolbe, 1895
81. Orphnus sudanicus Paulian, 1948
82. Orphnus tchadensis Benderitter, 1913
83. Orphnus thoracicus Linell, 1896
84. Orphnus tinantae Paulian, 1948
85. Orphnus transvaalensis Frolov, 2009
86. Orphnus tristis Pic, 1928
87. Orphnus usambaricus Petrovitz, 1971
88. Orphnus viduae Petrovitz, 1971

Subgenus Pachyorphnus Paulian, 1948
Type species: Orphnus subfoveatus Fairmaire, 1898, by original designation.

89. Orphnus clavipes Petrovitz, 1971
90. Orphnus dewittei Petrovitz, 1971
91. Orphnus subfoveatus Fairmaire, 1898

Subgenus Phornus Paulian, 1948
Type species: Orphnus strangulatus Paulian, 1948, by original designation.

92. Orphnus ferrierei Frolov et Akhmetova, 2016
93. Orphnus giganteus Paulian, 1948
94. Orphnus parastrangulatus Frolov et Akhmetova, 2016
95. Orphnus strangulatus Paulian, 1948
96. Orphnus renaudi Frolov et Akhmetova, 2016
97. Orphnus valeriae Frolov et Akhmetova, 2016

Subgenus Ronphus Paulian, 1948
Type species: Orphnus livingstonei Paulian, 1948, by original designation.

98. Orphnus demeyeri Frolov, 2013
99. Orphnus livingstonei Paulian, 1948
100. Orphnus massarti Paulian, 1948
101. Orphnus peringueyi Paulian, 1948
102. Orphnus planicollis Petrovitz, 1971
103. Orphnus quadrigibbosus Petrovitz, 1971
104. Orphnus simonii Petrovitz, 1971
105. Orphnus testaceus Paulian, 1948

Craniorphnus Kolbe 1895

Type species: Craniorphnus grandiceps Kolbe, 1895, by monotypy.

1. Craniorphnus grandiceps Kolbe, 1895

Goniorphnus Arrow, 1911

Type species: Goniorphnus felschei Arrow, 1911, by monotypy.

1. Goniorphnus felschei Arrow, 1911

Hybaloides Quedenfeldt, 1884

Type species: Hybaloides foveolatus Quedenfeldt, 1884, by monotypy.

1. Hybaloides foveolatus Quedenfeldt, 1884

Pseudorphnus Benderitter, 1913

Type species: Orphnus coquerelii Fairmaire, 1868, by monotypy.

1. Pseudorphnus carinatus Frolov, 2011
2. Pseudorphnus coquerelii (Fairmaire, 1868)
3. Pseudorphnus hiboni Paulian, 1959
4. Pseudorphnus olsoufieffi Paulian, 1977

Madecorphnus Paulian, 1992

Type species: Drepanognathus falciger Lansberge, 1886, original designation.

1. Madecorphnus aquilonius Frolov, 2012
2. Madecorphnus barclayi Frolov, 2012
3. Madecorphnus brunneus Frolov, 2010
4. Madecorphnus cuccodoroi Frolov, 2011
5. Madecorphnus dentatus Frolov, 2010
6. Madecorphnus falcatus Paulian, 1992
7. Madecorphnus falciger (Lansberge, 1886)
8. Madecorphnus falculoides (Paulian, 1977)
9. Madecorphnus fisheri Frolov, 2014
10. Madecorphnus grandis Frolov, Smith et Akhmetova, 2017
11. Madecorphnus hanskii Frolov, 2012
12. Madecorphnus montreuili Frolov, 2010
13. Madecorphnus niger Frolov, 2010
14. Madecorphnus pauliani Frolov, 2010
15. Madecorphnus perinetensis Frolov, 2010
16. Madecorphnus peyrierasi Frolov, 2010
17. Madecorphnus punctatus Frolov, 2010
18. Madecorphnus saintemariensis Frolov, 2014
19. Madecorphnus simplex Frolov, 2010
20. Madecorphnus tuberculatus Frolov, 2012

Triodontus Westwood, 1845

syn. Orphnidius (Benderitter, 1913)

Type species: Orphnus nitidulus Guérin-Méneville, 1844, by monotypy.

1. Triodontus alticola Paulian, 1977
    syn. Triodontus maroantsetrae Paulian, 1977. Synonymy by Frolov et al. (2016)
2. Triodontus ankarafantsikae Frolov, Montreuil et Akhmetova, 2016
3. Triodontus bicavatus (Fairmaire, 1905)
4. Triodontus copridoides Paulian, 1977
5. Triodontus fairmairei Frolov, Montreuil et Akhmetova, 2016
6. Triodontus hanskii Frolov, 2010
7. Triodontus hildebrandtii (Fairmaire, 1883)
8. Triodontus hova (Fairmaire, 1868)
    syn. Orphnus nigrita Brancsik, 1893. Synonymy by Frolov et al. (2016)
9. Triodontus inexpectatus Frolov, Montreuil et Akhmetova, 2016
10. Triodontus itremoi Paulian, 1977
11. Triodontus lemoulti Frolov, Montreuil et Akhmetova, 2016
12. Triodontus modestus (Benderitter, 1914)
13. Triodontus nitidulus (Guérin-Méneville, 1844)
     syn. Orphnus obsoletus Brancsik, 1893. Synonymy by Frolov et al. (2016)
     syn. Triodontus occidentalis Paulian, 1977. Synonymy by Frolov et al. (2016)
14. Triodontus owas Westwood, 1852
     syn. Triodontus vadoni Paulian, 1977. Synonymy by Frolov et al. (2016)
     syn. Triodontus perrotorum Paulian, 1977. Synonymy by Frolov et al. (2016)
15. Triodontus viettei Frolov, Montreuil et Akhmetova, 2016

Renorphnus Frolov et Montreuil, 2009

Type species: Orphnus clementi Petrovitz, 1971, by monotypy.

1. Renorphnus clementi (Petrovitz, 1971)

Tribe AEGIDIINI Paulian, 1984

Type genus: Aegidium Westwood, 1845

Aegidium Westwood, 1845

Type species: Aegidium colombianum Westwood.

1. Aegidium atlanticum Frolov, Grossi, & Vaz-de-Mello, 2015
2. Aegidium angustum Frolov, Akhmetova & Vaz-de-Mello, 2017
3. Aegidium bolivianum Frolov & Akhmetova, 2020
4. Aegidium bordoni Frolov, Akhmetova & Vaz-de-Mello, 2017
5. Aegidium borrei Paulian, 1984
6. Aegidium chalumeaui Rojkoff et Frolov, 2017
7. Aegidium colombianum Westwood, 1845
    syn. Aegidium elongatum Paulian, 1984, synonymy by Frolov et al. (2017).
8. Aegidium cribratum Bates, 1887
    syn. Aegidium asperatum Preudhomme de Borre, 1886, synonymy by Frolov et al. (2017).
    syn. Aegidium squamatum Bates, 1887: synonymy by Frolov et al. (2017).
9. Aegidium dierkensi Rojkoff et Frolov, 2017
10. Aegidium dominicense Cartwright and Chalumeau, 1977
11. Aegidium geayi Paulian, 1984
12. Aegidium gilli Frolov, Akhmetova & Vaz-de-Mello, 2017
13. Aegidium hostilioi Gasca-Álvarez and Deloya, 2020
14. Aegidium howdeni Frolov, Akhmetova & Vaz-de-Mello, 2017
15. Aegidium minor Paulian, 1984
16. Aegidium parvulus Westwood 1845
17. Aegidium onorei Frolov, Akhmetova & Vaz-de-Mello, 2017
18. Aegidium peresfilhoi Frolov, Akhmetova & Vaz-de-Mello, 2017
19. Aegidium peruanum Frolov, Akhmetova & Vaz-de-Mello, 2017
20. Aegidium rafaeli Frolov, Akhmetova & Vaz-de-Mello, 2017
21. Aegidium reichei Preudhomme de Borre, 1886
22. Aegidium sinuatum Frolov, Akhmetova & Vaz-de-Mello, 2017
23. Aegidium varians Frolov, Akhmetova & Vaz-de-Mello, 2017
24. Aegidium vincentiae Arrow, 1903

Aegidiellus Paulian, 1984

Type species: Phileurus alatus Laporte de Castelnau, by monotypy.

1. Aegidiellus alatus (Laporte de Castelnau, 1840)
    syn. Aegidium haedulus Westwood, 1845
2. Aegidiellus dentipenis Frolov, Akhmetova et Vaz-de-Mello, 2017
3. Aegidiellus zezaoi Frolov, Akhmetova et Vaz-de-Mello, 2017

Aegidinus Arrow, 1904

Type species: Aegidium guianensis Westwood, designated by Paulian (1984).

1. Aegidinus brasiliensis Arrow, 1904
2. Aegidinus candezei (Preudhomme de Borre, 1886)
    syn. Aegidinus sunidigea Colby, 2009
3. Aegidinus colbyae Frolov, Akhmetova & Vaz-de-Mello, 2019
4. Aegidinus cornutus Colby, 2009
    syn. Aegidinus unicus Colby, 2009
5. Aegidinus guianensis (Westwood, 1845)
    syn. Aegidinus crypticus Colby, 2009
    syn. Aegidinus oreibates Colby, 2009
6. Aegidinus howdenorum Colby, 2009
7. Aegidinus howeae Colby, 2009
8. Aegidinus noriegai Frolov, Akhmetova & Vaz-de-Mello, 2019
9. Aegidinus petrovi Colby, 2009
10. Aegidinus simulatus Colby, 2009
11. Aegidinus teamscaraborum Colby, 2009
12. Aegidinus steinheili (Harold, 1880)
13. Aegidinus tricornis Colby, 2009
14. Aegidinus venezuelensis Colby, 2009

Onorius Frolov et Vaz-de-Mello, 2015

Type species: Onorius inexpectatus Frolov et Vaz-de-Mello, 2015, by monotypy.

1. Onorius inexpectatus Frolov et Vaz-de-Mello, 2015
2. Onorius sp.

Paraegidium Vulcano, Pereira, et Martínez, 1966

Type species: Paraegidium costalimai Vulcano, Pereira, et Martínez, by monotypy.

1. Paraegidium barretoi Frolov, Akhmetova & Vaz-de-Mello, 2017
2. Paraegidium costalimai Vulcano, Pereira, et Martínez, 1966
3. Paraegidium howdeni Frolov, Akhmetova & Vaz-de-Mello, 2017
4. Paraegidium monneorum Frolov, Akhmetova & Vaz-de-Mello, 2017
5. Paraegidium oliveirai Frolov, Akhmetova & Vaz-de-Mello, 2017
6. Paraegidium pokornyi Frolov, Akhmetova & Vaz-de-Mello, 2017

Orphninae incertae sedis

Stenosternus Karsch, 1881

Type species: Stenosternus costatus Karsch, 1881, by monotypy.

1. Stenosternus costatus Karsch, 1881